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Since all sapient species are reproductively compatible genes work a little differently based upon the biology of the parents and how close or far away they are from each other in evolution.
It's best to think of it on this spectrum.
At the top, we have Darkborn, the first race.
Next, the Itru tribes who are the least different from the Darkborn physically.
Then the Platefins who have additional horn, and tail mutations.
Next the Thornscales, who begin to lose their tails and have further softened horns.
Then the oldest Davran species the Skins who have no fur with a notable percentage having vestigial tails.
Then the Longcoat who seemed to evolve sometime after the planet gained its rotation and shed some heat.
And finally, the Short-hairs, who (with shorter coats) were able to move more freely between warm and cold regions.
Then separately there are the two hybrid races of Featherlings and Tucktails.
If the parents differ in race then the child has up to an 85% chance of being hybridized. This is the maximum chance based upon parents on either side of the spectrum (Darkborn and Shorthair.) Chances are highest when the mother is the more evolved of the pair.
The only pairing with a higher than 85% chance of hybridization is one where both parents are already hybridized. If only one parent is hybridized, the child has a 50/50 shot of being a hybrid.
The grandfather gene is actually just another side of the hybrid gene. It causes a near-twin copy (both genetically. and physically speaking) of a parent's genes to transfer onto the child. It's called the grandfather because sometimes it can skip a generation. Generation twins are most common in Davran family lines and near nonexistent in pure Darkborn family lines.
It is proven that the grandfather gene and the hybrid gene are the same in family lines that practiced mixing Tucktail and Feathering parents. The child will almost always be a male generation twin of the mother or a female generation twin of the father. And if not, the child is still most likely a generation twin for a grandparent of the same gender. If no hybrid grandparent exists, but the child is not a "perfect" generation twin, then the eye color is inherited from one grandparent.
In accordance with its namesake, there is an inherent 5% chance of producing nonhybrid offspring between mismatched parents of all races that still differs from both. This only occurs if both parents also possess a mismatched parental pair with one member of the same race. For example, If a Tuckaill male has a Shorthair mother and he produces offspring with a Featherling woman who had a Shorthair father, there is an up to a 5% chance of the child being a pure-blooded shorthair as well as a generation twin. This also seems to follow the pattern of evolutionary chance, meaning it has the highest chance of happening in the Davrani family lines, reducing down the spectrum towards having a next to nonexistent chance of producing a Darkborn.