Technological achievement
The Krigan manage to turn the tide of war...at great cost. Their Queens, modified with Xavathun technology, can now produce tens of thousands of warriors and drone a day. Within months, the Empire has recuperated all previous losses. Within a few years, Krigan numbers swell a thousand-fold. But these new creatures are not fully Krigan. They are drones, bred only for war. With the Queens reduced to mindless breeding machines, the Primarchs take over leadership of the Empire.