Conductor Lizards and Subsequent Descendants

The conductor lizard is a fascinating group dating back around 40 million years. It has a strange genetic mutation that allows it to manage and even metabolize large amounts of electricity, including lightning. Living in rainforests, cloud forests and mangrove swamps on the Kalani island chain, they didn't have many natural predators, coming from an ancient lineage and originally evolving in a period in which the islands were relatively uncolonized by other species. Thus, this development lead to them seeking a new niche, and climbing treetops to try and harvest the raw energy they could absorb into their systems.
A new species of these lizards was formed when some began developing wings, much akin the gliding lizard of West Aguan. from specialized ribs with membranes stretched between them. These were developed not only to escape predators but also to take advantage of a potential new strategy, flying in packs higher than their surroundings to attract lightning and share it between them. This, however, was not as surefire as simply staying in a tall tree during the storm. So they got larger, and diversified into niches that did not previously exist, by adapting their wings, through turning their already segmented rib-wings into jointed structures.
Many of these lizards fused parts of these rib-spines, creating structures which allowed both control and agility and an ability to lock the wings in place for long flights. These proto-limbs allowed for a mass diversification into a multitude of large, migratory forms. A lot of the range of motion in their forelegs was sacrificed for these proto-limbs and the flight they offered. These adaptations were coupled with a growth in size to attract more lightning, and a general change in metabolic strategy. Growing larger, they had to support a higher calorie need (as well as a need for nutrients and minerals), and so could not fully subsist off lightning. As such they formed into a sort of food chain, with smaller gliders that hunt/scavenge birds, fish and insects, which in turn are preyed upon by larger dragonets.
To fly they sacrificed a lot of weight, be that organs, bone density, or muscle, and some specimens even developed internal gas-bags to aid with buoyancy, just enough to not sacrifice a measure of speed, simply creating a lighter form. These became the Great Dragons, great 6-limbed flying lizards with beardlike tendrils and large bodies which, though looking incredibly heavy, were actually quite light for the creature's size. Humans are known to have come in contact with them, especially in the temperate reaches around the Avarastinine sea, as they often seasonally make landfall and during this period can be quite territorial. Thus are they known as dragons, and they have been woven into myths and even religions all throughout the south. People who behold them usually do not make biological distinctions between them and the 4-limbed dragons of the equatorial regions, certainly ancient peoples never did.
Only the eldest in the conductor lizard lineages lack the defensive mechanism which allows them to set aside a portion of the electricity gathered to a specialized organ called the Patall, lined and shaped specially to keep the energy from decaying over extended periods of time. Not only does the patall create a sort of emergency store for these lizards but they can also actually eject the energy as a defence mechanism. This adaptation will be expanded upon by conductor lizard descendants, we'll get to that soon.
Ecologically these lizards have not affected their local ecosystems too much - they eat, including hunting and grazing, though that typically is outstripped by their dependence on lightning to deliver them their energy. Eating provides them with essential minerals and nutrients that they cannot get from lightning, however, so though it's relegated to a far smaller role it still is done, especially in dryer parts of the year. The southern wind dragons use their patalls in hunting, the electricity stored is actually converted to combustive energy along with the help of stored gas and other elements, creating jets of flame that can boil shoals of fish swimming near the surface alive. These flames are also used by males in mating displays when they make landfall along the Leasian moors and mounts.
Seabirds are known to opportunistically scavenge the hunts of Dragons and Dragonets, swooping in in hordes after a gout of fire leaves steam on the waves, and have developed an intestinal resistance to hot, boiled fish. Dragons and dragonets also hunt on land, but this is both infrequent and extremely hard to counter evolutionarily. As dragons are a seasonal predator and do not noticeably affect local wildlife populations, local species have not really changed physically. They have, however, changed both instincts and migration patterns, those instincts warding against large airborne objects and telling the deer to run into the cover of trees. There's not much you can do against a stream of fire or lightning so these instinctual adaptations have been the only ones.